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在未来，南极的上空出现了被称为“通路”的蘑菇云般的时空通路，外星生物“迦姆”利用它发动了对地球的攻击。经过33年惨烈的战斗，人类终于将“迦姆”封印于道路的彼端—

Plot Summary:Thirty-three years ago, an alien force known as the "JAM" invaded Earth through a dimensional portal over Antarctica. Earth's forces managed to drive the JAM away to a distant planet designated as "Fairy". While the majority of Earth's population is unaware of the JAM's presence, the war continues on Fairy, where Rei Fukai is an SAF (Special Air Force) pilot assigned to pilot "Yukikaze"-an advanced reconnaissance fighter plane equipped with a near-sentient A.I. that detects the presence of the JAM within its path.